List of Biosensors by Target

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When citing this resource, please reference the associated review paper [Pubmed]

Biosensor variant name links to biosensor details, year links to reference details.
TargetReadout MechVariants
Acetylation/Methylation
DNA methylationInten (☀)
DNA methylation BiAD☀DNA methylation BiAD sensor(2017) 
H3K9me3Inten (☀)
H3K9me3 BiAD☀H3K9me3 BiAD sensor(2017) 
Cell Cycle
Cell Cycle PhasesInten (☀)
FLIM (♣)
Fucci☀TNNT2-FUCCI(2022) ♣FUCCI-Red(2021) ☀Ki67p-FUCCI(2020) ☀Fucci(CA)2.1(2017) ☀NIR cell cycle reporter(2016) ☀NIR Fucci(2016) ☀Fucci4(2016) ☀Fucci2.1(2013) ☀Fucci2(2011) ☀zFucci(2009) ☀Fucci(2008) ☀S/G2/M-X(NC)(2008) 
G1 PhaseTransloc. (⛖)
G1 Phase Biosensor⛖G1 Phase Biosensor(2009) 
S PhaseInten (☀)
Transloc. (⛖)
S Phase Biosensor☀PCNA-mRuby(2017) ⛖S Phase Biosensor(2009) 
Cell Environment
Mechanical StrainFRET (⚡)
Ex. Rat.(★)
HP35⚡HP35-TS(2015) ⚡HP35st-TS(2015) 
PriSSM★PriSSM(2008) 
stFRET⚡cpstFRET(2012) ⚡sstFRET(2011) ⚡stFRET(2008) 
TSMod⚡FL-TSM(2017) ⚡VinTS(2016) ⚡TSMod F25 (2016) ⚡VinTL(2016) ⚡TSMod(2010) 
Molecular crowdingFRET (⚡)
CrGE⚡CRONOS(2021) ⚡CrGE2.3(2020) ⚡CrGE(2015) 
Crowding sensor⚡Crowding sensor(2015) 
pHEm. Rat. (☆)
Rat. (◧)
FRET (⚡)
Inten (☀)
FLIM (♣)
Ex. Rat.(★)
deGFP☆deGFP family(2002) 
Dendra◧Dendra2(2017) 
Dual pH and Cl sensor◧ClopHensor(2010) 
E^xGFP◧E^1GFP(2007) ◧E^2GFP(2006) 
FluBpH⚡FluBpH 5.7(2017) ⚡FluBpH 6.1(2017) ⚡FluBpH 7.5(2017) 
mtAlpHi☀mtAlpHi(2004) 
Native FP pH Sensitivity♣pHLIM(2022) ☆FIRE-pHLy(2021) ⚡pH-Lemon(2019) ☀mNect(2009) ☀pHVenus(2007) ⚡CFP-YFP pH FRET(2004) ☀EGFP(1998) ☀EYFP(1998) 
pHaROS★pHaROS(2019) ★pHaROS-red(2019) ★GRX1-pHaROS(2019) 
pHCEC◧pHCECSensor01(2008) 
pHluorins★(RpH)-LAMP1-3xFLAG(2020) ⚡pH-Lemon(2019) ★ER-sfpHluorin(2018) ★pHluorin2(2011) ☀Superecliptic-pHluorin(2000) ☀Ecliptic-pHluorin(1998) ★ratiometric-pHluorin(1998) 
pHRed★mCherryEA(2018) ★pHRed(2011) 
pHTomato☀syHTomato(2012) 
pHuji☀pHuji(2014) 
pHusion◧pHusion(2012) 
SRpHi◧SRpHi1(2017) ◧SRpHi2(2017) ◧SRpHi3(2017) 
SypHer★SypHer3s(2018) ★SypHer-2(2015) ☀SypHer(2011) 
XFpH⚡GFpH(2001) ⚡YFpH(2001) 
TemperatureFRET (⚡)
Rat. (◧)
Inten (☀)
ELP⚡ELP-TEMP(2021) 
gTEMP◧B-gTEMP(2022) ◧gTEMP(2017) 
mRFP☀mRFP1(2015) 
tsGFP☀tsGFP1(2013) ☀tsGFP2(2013) 
V+Inten (☀)
FRET (⚡)
BRET (◓)
Rat. (◧)
ArcLight Species Variants☀Frog Q174 ArcLight(2013) ☀Chicken Q175 ArcLight(2013) ☀Zebrafish Q175 ArcLight(2013) ☀Human Q193 ArcLight (2013) 
Ci-VSP based sensors☀ASAP4b(2023) ☀ASAP4e(2023) ⚡nirButterfly(2020) ☀Red-GEVIs(2019) ☀ASAP3(2019) ☀Marina(2017) ◓LOTUS-V(2017) ☀VSD-FR189-188(2017) ◧tdFlicR-VK-ASAP(2016) ☀tdFlicR1 delta 110AR(2016) ☀ASAP2f(2016) ☀FlicR1(2016) ☀FlicGR1(2016) ⚡Nabi2.213(2015) ☀Bongwoori(2015) ☀ArcLightning(2015) ⚡Chimeric VSFP-Butterfly CY(2014) ⚡Chimeric VSFP-Butterfly YR(2014) ☀ASAP1(2014) ⚡Mermaid2(2013) ⚡VSFP-CR(2012) ☀ArcLight A242(2012) ☀ElectricPk(2012) ⚡Chimera Cx(2012) ⚡VSFP Butterfly 1.2(2012) ⚡VSFP2.3(2010) ⚡VSFP2.42(2010) ⚡VSFP2.4(2009) ⚡Mermaid(2008) ☀VSFP3.1(2008) ⚡VSFP2.1(2007) 
Danio VSP based sensor☀Zahra 1 SE (227D) series(2015) ☀Zahara 2 SE (227D)(2015) ⚡Zahra 2(2012) ⚡Zahra 1(2012) 
Kv Based V sensors☀FlaSh IR(2002) ☀FlaSh L366A(2002) ⚡FlaSH (CFP) + FlaSh (YFP)(2002) ⚡VSFP1(2001) ☀FlaSh(1997) 
Na Channel based V Sensors☀SPARC(2002) 
Nematostella VSP based sensors⚡Nema(2012) 
rhodopsin based V sensors☀QuasAr6b(2023) ☀QuasAR6a(2023) ⚡Positron(2020) ☀Voltron(2019) ☀Archon2(2018) ☀VARNAM(2018) ☀Archon1(2018) ◧vArch1.0(2016) ☀FRET-opsin Ace2N-mNeon(2015) ☀FRET-opsin Ace1Q-mNeon(2015) ☀QuasAr2(2014) ☀QuasAr1(2014) ☀Archer1(2014) ⚡eFRET GEVI(2014) ☀MacQ-mCitrine(2014) ☀MacQ-mOrange2(2014) ☀FRET-opsin Mac-mCitrine(2014) ☀Arch-EEx variants(2013) ☀PROPS(2011) ☀Arch(D95N)(2011) 
Cellular Analytes
2OGInten (☀)
mOGsor☀mOGsor(2014) 
ADPFRET (⚡)
ADPrime⚡ADPrime(2019) 
AmmoniumRat. (◧)
Inten (☀)
AmTrac◧AmTryoshika1;3-LS-F138I -T78H(2017) ◧AmTryoshika1;3-LS-F138I(2017) ☀AmTrac(2013) 
ArabinoseFRET (⚡)
FLIPara⚡FLIPara-250n(2008) 
ArgFRET (⚡)
FRET Arg Reporter⚡FRET Arg Reporter(2007) 
ATPBRET (◓)
FRET (⚡)
Inten (☀)
Biolum. (⚫)
Ex. Rat.(★)
ATeam◓ARSeNL-ATP(2019) ◓BTeam(2016) ⚡ATeam1.03NL(2013) ⚡GO-Ateam(2011) ⚡ATeam1.03(2009) ⚡ATeam3.10(2009) 
EAF-based ATP biosensor⚡EAF-ATP(2013) 
iATPSnFR☀iATPSnFR2(2024) ☀iATPSnFR1.0(2019) ☀iATPSnFR1.1(2019) 
MaLion☀MaLionG(2018) ☀MaLionR(2018) ☀MaLionB(2018) 
Nano-lantern ATP⚫Nano-lantern ATP(2012) 
QUEEN★QUEEN-7mu(2014) ★QUEEN-2m(2014) 
Syn-ATP⚫Syn-ATP(2014) 
ATP:ADP RatioEx. Rat.(★)
Perceval★PercevalHR(2013) ★Perceval(2009) 
BCAAFRET (⚡)
OLIVe⚡OLIVe(2019) 
BDNFFRET (⚡)
Bescell⚡Bescell(2008) 
cAMPInten (☀)
FRET (⚡)
BRET (◓)
Rat. (◧)
Biolum. (⚫)
cADDis☀cADDis-green(2016) 
camps⚡cAMPFIRE-L(2022) ⚡cAMPFIRE-M(2022) ⚡cAMPFIRE-H(2022) ⚡Epac2-camps300-cit(2010) ⚡Epac2-camps300(2009) ⚡HCN2-camps(2006) ⚡PKA-camps(2004) ⚡Epac1-camps(2004) ⚡Epac2-camps(2004) 
CAMYEL◓CAMYEL(2007) 
CUTie⚡CUTie(2017) 
Epac-S⚡Epac-S H187(2015) ⚡Epac-S H188(2015) ⚡H96(2011) ⚡H74(2011) ⚡H81: GFP(nd)-EPAC(dDEP)-mRFP (2008) ⚡H90: CFP(nd)-EPAC(dDEP/CD)-cp173Venus(d)(2008) ⚡H84: CFP(nd)-EPAC(dDEP/CD)-Venus(d)(2008) 
Epac1 (delta DEP-CD) based cAMP biosensors⚡CEPAC*(2011) ⚡CFP-Epac(delta DEP-CD)-YFP(2004) 
Flamindo☀Pink Flamindo(2017) ☀Flamindo2(2014) ☀Flamindo(2013) 
FPX◧pPHT-PKA(2015) 
ICUE⚡GR-ICUE2(2020) ☀RAB-ICUE(2018) ⚡ICUE-YR(2011) ⚡ICUPID(2011) ⚡ICUE3(2009) ⚡ICUE2(2008) ◓Rluc-EPAC-YFP(2008) ⚡ICUE1(2004) ⚡xpkLoRel(1967) 
mCRIS based cAMP biosensor⚡cit-mCNBD-cer(2013) 
mICNBD☀DDcAMP(2022) ☀G-Flamp1(2022) ☀G-Flamp2(2022) ☀G-Flamp2b(2022) 
mICNBD-FRET⚡mICNBD-FRET(2016) 
Nano-lantern cAMP⚫Nano-lantern cAMP1.6(2012) 
R-FlincA☀R-FlincA(2018) 
R1alpha #7⚡R1alpha #7(2016) 
Split Luc cAMP biosensor⚫22F(2011) ⚫Split Luc cAMP biosensor(2008) 
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
xpkLoRel
YFP-PKAc + CFP-PKAr◓R1-Rluc + GFP-C(2006) ◓R2-Rluc + GFP-C(2006) ⚡delta PKA RIIb-CFP + PKAc-YFP(2006) ⚡YFP-PKAc + CFP-PKAr(R230K)(2004) ⚡YFP-PKAc + CFP-PKAr(2002) ⚡PKAc-S65T + PKArII-EBFP(2000) 
cGMPFRET (⚡)
Inten (☀)
Rat. (◧)
Ex. Rat.(★)
cGES⚡ThPDE5VV(2019) ☀Cygnus(2010) ⚡cGES-DE5(2006) 
cGKI based cGMP sensors⚡cGi family(2007) 
cGull☀Green-cGull(2017) 
CGY⚡CGY-del1(2000) 
CUTie⚡CUTie2(2021) 
cyGNAL◧cyGNAL(2019) 
cygnet⚡cygnet-2.1(2005) ⚡cygnet-2(2001) 
FlincG⚡FGA^M(2013) ☀H6-FGB(2013) ★delta-FlincG(2008) 
PfPKG⚡PfPKG Yellow(2019) ⚡Red PfPKG(2019) 
CitrateFRET (⚡)
FLIP-Cit⚡FLIP-Cit-Y(2011) 
COInten (☀)
COSer☀COSer(2012) 
F-1,6-BPFRET (⚡)
Inten (☀)
fructose‐1,6‐bisphosphate⚡HYlight(2022) ☀Glycolytic Flux Sensor(2019) 
GlnFRET (⚡)
FLIPQ⚡FLIPQ-TV3.0(2012) 
GlucoseFRET (⚡)
Ex. Rat.(★)
Inten (☀)
FLIPglu⚡FLII^12Pglu-Y Series(2008) ⚡FLIPglu-Y delta 13 (2006) ⚡FLII^XPglu-Y Series (2005) ⚡FLIPglu-600u-delta(X) series(2005) ⚡FLIPglu-Y series(2003) 
GGBP★FGBP(2018) 
GIP⚡AcGFP1-GBPcys-mCherry(2010) ⚡GIP C_0Y_i(2008) ⚡GIP(2003) 
Glifon☀Red Glifon(2022) ☀Green Glifon(2019) 
GTPEx. Rat.(★)
GEVAL★GEVAL30(2017) ★GEVAL260(2017) ★GEVAL530(2017) ★GEVAL1150(2017) ★GEVAL2300(2017) 
HemeFRET (⚡)
Rat. (◧)
Inten (☀)
CHY⚡CH49Y(2017) 
CISDY⚡CISDY-9(2015) 
Fluorescence quenching Heme◧HS1(2016) ◧HS1-M7A(2016) ☀CG6(2012) 
HisRat. (◧)
FRET (⚡)
FHisJ◧FHisJ(2017) 
FLIP-HisJ⚡FLIP-cpHisJ194(2009) 
InsulinRat. (◧)
RINS◧RINS1(2017) 
LactateFRET (⚡)
Laconic⚡Laconic(2013) 
LCACoAsRat. (◧)
LACSer sensor◧LACSer(2021) 
maltoseBRET (◓)
FRET (⚡)
Inten (☀)
FLIPmal◓GFP2-MBP-RLuc2(2013) ⚡FLIPmal-Y delta1(2008) ⚡FLIPmal-Y Series(2002) 
MBP☀PPYF-green(2011) 
NAD+Inten (☀)
FRET (⚡)
NAD+ Biosensor☀NAD+ Biosensor(2016) 
NAD-Snifit⚡NAD-Snifit(2018) 
NAD+/NADH RatioRat. (◧)
Ex. Rat.(★)
Inten (☀)
Peredox◧Peredox(2011) 
SoNar★SoNar(2015) 
T-Rex Based☀RexYFP(2014) 
NADHInten (☀)
Frex☀Frex(2011) ☀FrexH(2011) 
O2Inten (☀)
FRET (⚡)
dUnOHR hypoxia-reoxygenation sensor☀dUnOHR hypoxia-reoxygenation sensor(2016) 
FluBO⚡FluBO(2012) 
PnInten (☀)
EcPhnD based☀EcPhnD-cpGFP(2011) 
PyruvateInten (☀)
FRET (⚡)
PdhR Based☀PyronicSF(2020) ⚡PYRATES(2017) ⚡Pyronic(2014) 
riboseFRET (⚡)
FLIPrib⚡FLIPrib-Y(2003) 
sucroseFRET (⚡)
FLIPsuc⚡FLIPsuc-Y(2006) 
TrehaloseFRET (⚡)
FLIP⚡FLIPSuc90μ∆1Venus(2016) 
TrpFRET (⚡)
Rat. (◧)
FLIPW⚡FLIPW(2007) 
GRIT◧GRIT(2023) 
Ions
As3+FRET (⚡)
SenALiB⚡SenALiB(2019) 
Ca2+FLIM (♣)
Inten (☀)
BRET (◓)
Biolum. (⚫)
FRET (⚡)
Em. Rat. (☆)
Rat. (◧)
Ex. Rat.(★)
Ca-FLITS♣Tq-Ca-FLITS(2021) 
Ca2+ Snapshot☀FLiCRE(2020) ☀Cal-Light(2017) ☀FLARE(2017) 
CaMBI◓Orange CaMBI 110(2019) ◓Orange CaMBI 300(2019) ⚫Blue CaMBI(2019) ◓Green CaMBI(2019) 
Cameleons♣4mtD3mC3(2019) ◓D3 BRET(2015) ⚡D1GO-Cam(2012) ⚡CaYin1(2011) ⚡CaYang1(2011) ⚡YC-Nano(2010) ⚡D3cpv(2006) ⚡D4cpV(2006) ⚡YC3.60(2004) ⚡D1(2004) ⚡YC2.12(2002) ⚡YC6.1(2001) ⚡YC2.1(1999) ⚡Cameleon 3(1997) ⚡Split-Cameleon(1997) 
Camgaroo☀Camgaroo2(2001) ☀Camgaroo1(1999) 
CaMPARI☀CaMPARI2(2018) ☀CaMPARI(2015) 
CASE☀Case12(2007) ☀Case16(2007) 
CatchER☀CatchER(2011) 
ddFP Ca2+ sensor☀ddRFP Ca2+ sensor(2012) 
ER-GCaMP☀ER-GCaMP6-210(2017) ☀ER-GCaMP6-150(2017) ☀ER-GCaMP3-373 / GCaMPer-10.19(2015) ☀CEPIA1er(2014) 
ER-GECO☀G-CEPIA1er(2014) ☀R-CEPIA1er(2014) ☆GEM-CEPIA1er(2014) 
FGCaMP☀FRCaMP(2020) ◧FGCaMP7(2020) ★FGCaMP(2017) 
FPX Ca2+ Sensors◧single polypeptide FPX Ca2+ sensor(2015) ◧tripartate FPX Ca2+ Sensor(2015) 
GAF-CaMP☀GAF-CaMP3-sfGFP(2020) ☀GAF-CaMP2(2019) ☀GAF-CaMP(2018) 
GAP☀GAP3(2016) ☀GAP1(2014) 
GCaMP☀jGCaMP8(2023) ☀GCaMP6s-Q(2022) ☀jYCaMP1(2020) ☀jYCaMP1s(2020) ☀GCaMP7(2019) ☀axon-GCaMP6(2018) ☀ncpGCaMP6s(2018) ◧MatryoshCaMP6s(2017) ◧sfMatryoshCaMP6s-T78H(2017) ◧sfMatryoshCaMP6s(2017) ◧GCaMP-R-3(2017) ◧GCaMP-R-6f(2017) ◧GCaMP-R-6s(2017) ☀GCaMP3fast(2015) ☀GCaMP3bright(2015) ☀GCaMP2.2Low(2014) ☀GCaMP6m(2013) ☀GCaMP6s(2013) ☀GCaMP6f(2013) ☀G-CaMP 3-8 (Nakai Variants)(2012) ☀GCaMP5G(2012) ☀GCaMP-HS(2011) ☀GCaMP3(2009) ☀GCaMP2(2006) ☀GCaMP(2001) 
GECO☀NEMO(2023) ☀NIR-GECO2(2020) ☀R-CEPIA4mt(2020) ☀R-CEPIA3mt(2020) ☀mNG-GECO1(2020) ☀NIR-GECO1(2019) ◓LUCI-GECO1(2018) ☀K-GECO1(2018) ☀jRGECO1a(2016) ★REX-GECO(2014) ☀LAR-GECO(2014) ☀R-GECO1.2(2013) ☀CAR-GECO(2013) ☀O-GECO(2013) ☀GR-GECO(2013) ☀G-GECO1.1(2011) ★GEX-GECO(2011) ☆GEM-GECO(2011) ☀R-GECO1(2011) ☀B-GECO(2011) ☀G-GECO1.2(2011) ☀G-GECO1(2011) 
GreenT-EC☀GreenT-EC(2023) 
Nano-lantern Ca2+◓CeNL(Ca2+)_110u(2018) ◓CeNL(Ca2+)_19u(2018) ◓GeNL(Ca2+)(2016) ◓CNL (Ca2+) (2015) ◓ONL (Ca2+)(2015) ⚫Nano-lantern Ca2+(2012) 
NCaMP☀NCaMP7(2020) 
NTnC☀YTnC(2018) ☀iYTnC2(2018) ☀NTnC(2016) 
pericam☀flash-pericam(2001) ☀inverse-pericam(2001) ◧ratiometric-pericam(2001) 
RCaMP☀jRCaMP1a(2016) ☀jRCaMP1b(2016) ☀R-CaMP2(2015) ☀RCaMP1h(2013) ☀R-CaMP1.07(2012) 
TN Ca2+ Sensors◓CalfluxVTN(2016) ⚡Twitch series(2014) ⚡RS-TN-XL(2013) ⚡YO-TnC1.0(2012) ⚡TN-XXL(2008) ⚡TN-XL(2006) ⚡TN-L15(2004) ⚡TN-humTnC(2004) 
Cd2+FRET (⚡)
Cd-FRET⚡Cd-FRET2(2011) 
Cu+Inten (☀)
FRET (⚡)
CreiLOV☀CreiLOV(2020) 
Yeast Copper regulator based⚡Ace1-FRET(2011) ⚡Mac1-FRET(2011) ⚡Amt1-FRET(2010) 
Halide IonsFRET (⚡)
Rat. (◧)
Inten (☀)
Clomeleon⚡SuperClomeleon(2013) ⚡Cl-sensor(2008) ⚡Clomeleon(2000) 
Dual pH and Cl sensor◧ER-LSSmsfClopHensor(2022) ◧LSSmClopHensor(2016) ◧ClopHensor(2010) 
Halide Sensitive Fluorescent Proteins☀mBeRFP-S94V/R205Y(2021) ☀GR1(2021) ☀mNeonGreen-R195Y(2019) ☀phiYFP(2019) ☀Cl-YFP(2014) ☀ClsM(2014) ☀YFP(H148Q, I152L)(2001) ☀YFP(H148Q)(2000) 
Hg(II)Inten (☀)
eGFP based Hg sensor☀eGFP205C(2008) 
IFP based Hg sensor☀IFP based Hg sensor(2011) 
K+FRET (⚡)
Inten (☀)
GEPII⚡KIRIN1-GR(2019) ⚡KIRIN1(2019) ⚡GEPII 1.0(2017) 
GINKO☀KRaION1(2022) ☀GINKO2(2022) ☀GINKO1(2019) 
Ln3+FRET (⚡)
LaMP⚡LaMP1(2019) 
Mg2+FRET (⚡)
Rat. (◧)
MagFRET⚡MARIO(2018) ⚡MagFRET-1(2013) 
MagIC◧MagIC(2015) 
Mn2+FRET (⚡)
MnLaMP⚡MnLaMP1(2022) ⚡MnLaMP2(2022) 
PhosphateFRET (⚡)
FLIPPi⚡FLIPPi-30m(2006) ⚡FLIPPi-5mu(2006) 
Zn2+BRET (◓)
FRET (⚡)
Inten (☀)
Rat. (◧)
Atox1 WD4 Based Zn2+ Probes◓BLCALWY-1(2016) ⚡redCALWY(2013) ⚡eCALWY series (2009) ⚡CALWY(2007) ⚡CFP-Atox1 + WD4-YFP(2006) 
GZnP☀GZnP3(2019) ☀GZnP2(2018) ☀GZnP1(2016) 
Minimal Zinc Finger Based Zn Probes☀FRISZ(2023) ◧ZifCV1.173(2019) ☀ZIBG2(2019) ☀ZnGreen2(2016) ⚡mito-ZifCY1.173(2012) ⚡Cys2Hys2 - Zn2+ Sensor(2009) ⚡His4-Zn2+ sensor(2009) 
Zap1 Based Zn Probes☀RZnP1(2022) ☀ZIBG1(2019) ⚡ZapCV2(2017) ⚡ZapCV5(2017) ☀ZnGreen1(2016) ⚡ZapCY1(2011) ⚡ZapCY2(2011) ⚡ZF1/2-FRET(2006) 
ZinCh◓BLZinCh-2(2016) ◓BLZinCh-3(2016) ⚡eZinCh-2(2015) ⚡eZinCh-1(2011) ⚡Cly9-2His(2008) ⚡ZinCh-x(2007) 
Kinases/Phosphatases
AblFRET (⚡)
Abl Indicator⚡Abl Indicator(2001) 
AktFRET (⚡)
Transloc. (⛖)
Ex. Rat.(★)
FLIM (♣)
Akind⚡Akind(2006) 
Akt translocation Reporters⛖Akt-FoxO3a-KTR-EGFP(2016) ⛖FoxO1-clover(2015) 
AktAR★ExRai-AktAR2(2021) ⚡VstagAktAR(2020) ★ExRai-AktAR(2018) ⚡AktAR2(2015) ⚡AktAR(2008) 
AktUS⚡Aktus(2003) 
BKAR⚡BKAR v2(2014) ⚡BKAR(2005) 
Dual Labeled Akt⚡GFP-PKB-RFP(2007) ♣GFP-AKT-YFP(2003) 
Eevee-Akt⚡Eevee-iAkt(2014) ⚡Eevee-Akt(2011) 
ReAktion⚡ReAktion1(2007) 
AMPKFRET (⚡)
Ex. Rat.(★)
AMPfret⚡AMPfret 2.1(2019) 
AMPKAR⚡AMPKAR-EV(2017) ⚡bimABKAR(2015) ⚡ABKAR(2015) ⚡AMPKAR(2011) 
ExRai-AMPKAR★ExRai-AMPKAR(2022) 
ATM kinaseFRET (⚡)
ATOMIC⚡ATOMIC(2007) 
Aurora B kinaseFRET (⚡)
Aurora B sensor⚡Aurora B sensor (Chu)(2011) ⚡Aurora B Sensor (Fuller)(2008) 
Aurora Kinase AFLIM (♣)
FRET (⚡)
AURKA Biosensor♣ShadowG-AURKA-mTurquoise2(2019) ⚡GFP-AURKA-GFP(2019) ♣ShadowY-AURKA-mTurquoise2(2019) ⚡superYFP-AURKA-mTurquose2(2019) ⚡AURKA Biosensor(2016) 
B-RafFRET (⚡)
Prin-BRaf⚡Prin-BRaf(2006) 
Bcr-AblFRET (⚡)
Pickles⚡Pickles-2.3(2010) 
C-RafFRET (⚡)
Prin-CRaf⚡Prin-CRaf(2005) 
CaMKIIFLIM (♣)
FRET (⚡)
Camui♣Camui-mRmC(2016) ♣ShadowG-Camui(2015) ⚡Camui?-CR (Clover-mRuby2)(2012) ♣green-Camui (alpha)(2009) ♣mRFP/GFP-Camui(2008) ⚡Camui(2005) 
FRESCA⚡FRESCA(2019) 
K2alpha⚡YC-K2alpha(2013) ⚡RY-K2alpha(2013) ⚡RS-K2Alpha(2013) 
CaNFRET (⚡)
CaN Activation⚡RY-CaN(2013) ⚡YC-CaN(2013) 
CaNAR⚡CaNAR2(2014) ⚡CaNAR1(2008) 
CaNARi⚡CaNARi(2014) 
DuoCaN⚡DuoCaN(2015) ⚡UniCaN(2015) 
CDK1FRET (⚡)
CyclinB1-Cdk1 activity sensor⚡CyclinB1-Cdk1 activity sensor V2(2014) ⚡CyclinB1-Cdk1 activity sensor V1(2010) 
CDK2Transloc. (⛖)
CDK2 Translocation Reporter⛖DHB-Ven(2013) 
DAPK1FRET (⚡)
DAPK1 sensor⚡DAPK1(2011) 
EGFRFRET (⚡)
ECaus⚡ECaus(2008) 
EGFR Reporter⚡EGFR Reporter(2001) 
FLAME⚡FLAME(2004) ⚡PTB-EYFP, EGFR-ECFP(2004) 
Picchu-Z⚡Picchu-Z(2005) 
ERKFRET (⚡)
Inten (☀)
FLIM (♣)
BRET (◓)
Transloc. (⛖)
Biolum. (⚫)
FLINC (◀)
Rat. (◧)
EAS⚡EAS3(2005) 
EKAR⚡EKAREN5(2021) ⚡EKAREN4(2021) ⚡EKAR4(2020) ⚡fRR-EKARev(2020) ☀RAB-EKARev(2018) ⚡VV FLARE EKAR(2018) ⚡C3C3 FLARE EKAR(2018) ⚡mCh-mCh FLARE EKAR(2018) ⚡VcpV FLARE EKAR(2018) ♣EKARet(2017) ♣EKAR^dual(2017) ⚡bimEKAR(2015) ⚡EKAR3(2015) ⚡EKAREV-TVV(2014) ⚡EKAR-TVV(2014) ◓REV(2013) ⚡EKAR2G(2013) ⚡EKAREV(2011) ⚡EKAR(2008) 
ERK-KTR⛖ERK-KTR(2014) 
Erkus⚡Erkus(2007) 
ERKy⚡ERKy(2012) 
FIRE⚫CBGreen-FIRE(2020) ☀FIRE(2013) 
FLINC-EKAR1◀FLINC-EKAR1(2017) 
FPX ERK◧FPX EKAR(2015) 
Miu2⚡Miu2(2006) 
FAKFRET (⚡)
FAK Activation Biosensor⚡CYFAK413(2008) 
FAK Autophosphorylation Biosensor⚡FAK Autophosphorylation Biosensor(2008) 
FAK sensor⚡FAK sensor(2011) 
Fus3Transloc. (⛖)
Far1-SKARS⛖Far1-SKARS(2015) 
FynFRET (⚡)
FRET⚡Fyn-FRET(2019) 
saFRET⚡Fyn-saFRET (2021) 
GCKFRET (⚡)
GCK Activation Biosensors⚡FRET-GCK (cp173-mCer3/mVen)(2016) ⚡FRET-GCK (mCer/mVen)(2011) ⚡Cerulean-GCK-mCit(2004) ⚡FRET-GCK(2002) 
H3-S10pFRET (⚡)
H3S10p biosensor⚡H3S10p biosensor(2018) 
H3-S28pFRET (⚡)
Histone Phosphorylation reporter⚡Histone Phosphorylation reporter(2004) 
INSRBRET (◓)
FRET (⚡)
Inten (☀)
Insulin Receptor Activation BRET assay◓Insulin Receptor Activation BRET assay(2001) 
Phocus⚡Phocus2(2002) ⚡Phocus-2pp nes(2002) 
sinphos☀cyan-sinphos(2004) ☀green-sinphos(2004) ☀yellow-sinphos(2004) 
JNKFRET (⚡)
Transloc. (⛖)
dJUN-FRET⚡dJUN-FRET(2008) 
JNK-KTR⛖JNK-KTR(2014) 
JNKAR⚡NIR JNKAR(2018) ⚡bimJNKAR(2015) ⚡JNKAR1EV(2011) ⚡JNKAR1(2010) 
JuCKY⚡JuCKY(0) 
Kss1Transloc. (⛖)
Ste7-SKARS⛖Ste7-SKARS(2015) 
LATSBiolum. (⚫)
LATS-BS⚫LATS-BS(2018) 
LckFRET (⚡)
Lck Activation sensor⚡CLckY-2(2009) 
MAPK or MK2FRET (⚡)
GMB⚡GMB(2001) 
MARKFRET (⚡)
MARK sensor⚡MARK-AR1(2011) 
Mpk1Transloc. (⛖)
Mkk2-SKARS⛖Mkk2-SKARS(2015) 
mTORC1FRET (⚡)
TORCAR⚡TORCAR(2015) 
p38FRET (⚡)
Transloc. (⛖)
p38 activity reporter⚡p38 activity reporter(2015) 
p38-KTR⛖mKO-MK2(2018) ⛖p38-KTR(2014) 
PAK1FRET (⚡)
Pakabi⚡Pakabi(2009) 
PDGFRFRET (⚡)
PDGFR Biosensor⚡PDGFR Biosensor(2017) 
PDK1FRET (⚡)
PARE⚡PARE(2011) 
PKA